Doctoral Researcher in Experimental evolution and evolutionary genetics The Kronholm lab is looking to hire a doctoral researcher (PhD student position) to join a project combining experimental evolution and genetic analysis to study the properties of mutations that arise during adaptation. The genetics, epigenetics, and evolution group lead by Ilkka Kronholm studies the properties of mutations and spontaneous epigenetic changes, and genetic architecture of complex traits. Our fundamental interest is in the properties of mutations, and how do they interact. Our research questions are focused on understanding how different intrinsic and extrinsic factors affect evolutionary adaptation. As model systems we use fission yeast and the filamentous fungus Neurospora crassa. We are currently working on a large experimental evolution study using fission yeast, with two postdocs and one other PhD researcher working on different questions. This project is part of a larger project funded by the ERC. The idea with this project is that we will be exploring how do different mutations that have arisen during adaptation affect different life-history and physiological traits, and how those changes are translated in to effects on fitness: so investigating the genotype - phenotype - fitness map and possibly effects of epistatic interactions among the mutations. Our system allows very detailed investigation of the relationship between genotype and phenotype, since genetic variation is limited and we can leverage the full power of fission yeast genetics. Moreover, in this project the candidate also has some freedom to explore different questions, so that exact questions can also be tailored to the candidates interests, within the overall framework of the project. The doctoral researcher is expected to contribute to the current project, plan and perform research, write manuscripts and finally a PhD thesis. There is also a limited amount of teaching and other coursework that is part of the PhD studies. You are the person we are looking for if you have a master's degree in evolutionary biology, population genetics, genetics or a related discipline with a strong interest in evolutionary genetics. Previous experience with fungal genetics or microbiology is not required, but having some background in microbial work can be an advantage, as well as good computational skills and solid understanding of statistics, and in particular the R environment. Good written and oral communication skills in English are required. The position can start in beginning of January 2026 or somewhat later as agreed. There is going to funding for three years, and there are several options to apply for funding for the fourth year if needed. What does the University of Jyväskylä offer as an employer? At the University of Jyväskylä, you are a recognized member of our community with an ample opportunity to be drawn into international research. You get to participate in our international and multidisciplinary community, where everybody's welfare is essential. You will work in an inspiring and lively campus area and an environment that supports a healthy and active lifestyle. Finland has a high standard of living with healthcare, free schooling (also in English), affordable childcare, and good family benefits. The city of Jyväskylä is located in central Finland amidst Finnish lakes and has excellent opportunities for different nature, outdoor, and sports activities. It is a major educational center, whose large student population is responsible for a vibrant cultural scene. To find useful information about the University of Jyväskylä, the City of Jyväskylä, and living in Finland, see the University's International Staff Guide. The initial annual salary will be approximately 28,000 EUR (gross income, including a holiday bonus). With progress of the thesis work, the salary will be revised in accordance with the Collective Agreement of Finnish Universities. The employment starts with a trial period of six months. Please attach the following documents to your application: 1. Curriculum vitae, composed according to good scientific practice and considering the template for researcher´s curriculum vitae by The Finnish Advisory Board on Research Integrity (https://tenk.fi/en/advice-and-materials/template-researchers-curriculum-vitae) and include contact information of two people willing to provide a reference. 2. Cover letter explaining your motivation, research interests, and plans. Please do not use AI to write your cover letter. 3. Copy of your Master's degree or a statement explaining your schedule for obtaining it. Please submit your application at the latest by 31 October 2025 11:45 pm (GMT +2). Please do not submit your application by email, but use the online application form: https://ats.talentadore.com/apply/doctoral-researcher-in-evolutionary-genetics/Z93bKA or the same as a short link: https://tinyurl.com/mpp8rx7u For further details or questions, please contact: Associate Professor Ilkka Kronholm, ilkka.kronholm@jyu.fi. website: ikronholm.net Ilkka Kronholm Associate Professor of Evolutionary Biology Department of Biological and Environmental Science University of Jyväskylä ilkka.kronholm@jyu.fi +358 417491785 @kronholmlab "Kronholm, Ilkka" (to subscribe/unsubscribe the EvolDir send mail to golding@mcmaster.ca)