*PhD studentship in sex and tardigrade evolution* Fully funded 4-year PhD studentship in tardigrade evolution at the Jagiellonian University in Krakow (Poland) in the team of Prof. ukasz Michalczyk, in collaboration with Prof. Mark Blaxter (Wellcome Sanger Institute, Cambridge, UK). The main goal of the project is to take a unique advantage of applying cutting-edge methodology of genome sequencing to a fascinating model, tardigrades, in order to answer one of the fundamental questions in life sciences: why are some asexual lineages able to persist over long evolutionary time, whereas the majority of them seem to be evolutionary dead ends? Given that genetic recombination seems to be the key to evolutionary success, the prediction is that ancient asexual lineages exhibit genomic signatures of recombination, whereas most recent asexuals do not. In other words, the project will provide first large scale, systematic comparative analysis of the genomic architecture of ancient and recent asexuals. The PhD studentship presents a great and unique opportunity to answer one the fundamental questions in life sciences using a fascinating model and cutting edge methods. Importantly, the position also offers additional training in genomics and transcriptomics in one of the world's leading institutions in genomics, the Wellcome Sanger Institute in Cambridge, UK. Prof. Mark Blaxter, Head of the Tree of Life Programme at Sanger, is a formal and the key collaborator on the project. Although the evolution of sex and tardigrades are the central project aims, the system provides potential for a dedicated student to explore their own ideas on other topical tardigrade biology and evolutionary questions. A dedicated student will have a great chance not only to learn a range of transferable skills, but also to build a sound publication record which will help them to get an attractive PostDoc or industry position in the future. The successful candidate will be involved in will be involved in field work (sample collection), tardigrade identification, culturing, DNA extraction, genome & transcriptome sequencing and assembly, as well as in phylogeny reconstruction. The PhD Student will be also required to prepare first drafts of some manuscripts and will be involved in the promotion of results at seminars and conferences. The PhD programme in Biology at the Jagiellonian University is run entirely in English and it includes some obligatory and facultative classes. The programme is open to all nationalities and there are no tuition fees. The PhD stipend is 5  500  PLN per month (typically, cost of life in Krakow for a PhD student is ca. 4  000 PLN/month). Deadline for applications: 6th of June 2024. More information on the project, university, our team and the application procedure is available here: http://tardigrada.edu.pl/PhD.htm Prof. ukasz Michalczyk LM@tardigrada.net ukasz Michalczyk (to subscribe/unsubscribe the EvolDir send mail to golding@mcmaster.ca)