The 2025 Workshop on Molecular Evolution at the Marine Biological Laboratory in Woods Hole, MA will be held May 22 to June 1, 2025. Founded in 1988, the Workshop on Molecular Evolution is the longest-running workshop of its kind. The Workshop is the premier program for integrating the methods, theory, and applications of molecular phylogenetics, statistical genetics, molecular evolution, and related disciplines. Students work closely with internationally recognized scientists, receiving (i) high-level instruction in the principles of molecular evolution and phylogenetics; (ii) advanced training in statistical methods best suited to modern datasets and biological questions concerning species, populations, or infectious diseases; and (iii) hands-on experience with the latest software tools (often from the authors of the programs they are using). The material is delivered via lectures, discussions, and bioinformatic exercises motivated by contemporary topics in molecular evolution. Recently added topics to the course include deep-time phylogenomics, phylodynamics, and comparative methods. A hallmark of this workshop is the direct interaction between students and field-leading scientists. The workshop serves graduate students, postdocs, and established faculty from around the world seeking to apply the principles of molecular evolution to questions of both basic and applied biological sciences. A priority of this workshop is to foster an environment where students can learn from each other as well from the course faculty. As the course progresses, participants learn how to use the following software and tools to address questions concerning the origins, maintenance, and function of molecular variation: ASTRAL, BEST, FASTA, IQ-TREE, MIGRATE, MAFFT, MP-EST, Open Tree, RAxML, RevBayes, PAML, PAUP*, SNaQ, and SVD Quartets. Students will have the opportunity to work with software on their own laptops as well as receive training on how to use the same programs on a computer cluster. In 2025, the confirmed course instructors include Peter Beerli, Joseph Bielawski, Jeremy Brown, Belinda Chang, Scott Edwards, Laura Eme, Mandev Gill, Tracy Heath, John Huelsenbeck, Sungsik Kong, Lacey Knowles, Laura Kubatko, Paul Lewis, Emily Jane McTavish, Claudia Solis-Lemus, David Swofford, Anne Yoder, and Rosana Zenil-Ferguson. Deadline for applications is January 29, 2025: https://www.mbl.edu/education/advanced-research-training-courses/course-offerings/workshop-molecular-evolution More information on the Workshop is available on the course website: https://molevolworkshop.github.io For further information, please contact Workshop co-Directors: Tracy Heath and Jeremy Brown at moledirector@mbl.edu. Jeremy M Brown (to subscribe/unsubscribe the EvolDir send mail to golding@mcmaster.ca)