Michigan State U. LampreyEcotypeGenomics Description: We seek applications for a PhD assistantship focused on the genomic basis of parasitic and non-parasitic ecotypes in Ichthyomyzonlamprey. Nonparasitic lampreys have repeatedly evolved from parasitic species that feed on the blood of fish hosts. This shift in life history represents a loss of a complex trait, where nonparasitic lampreys transition directly from larvae into reproductive adults without developing the apparatus or appetite for parasitic feeding. The goal of this project is to understand the genomic and developmental basis of parasitism versus nonparasitism in the silver lamprey (I. unicuspis) and northern brook lamprey (I. fossor), two closely related ecotypes with ongoing gene flow in the Laurentian Great Lakes. The selected student will lead a collaborative project with two primary components: 1) a common garden study in which full reciprocal crosses of Iu and If will be raised at the USGS Hammond Bay Biological Station (HBBS; https://www.usgs.gov/centers/great-lakes-science-center/science/hammond-bay-biological-station) in Millersburg, MI and sampled for transcriptomic and phenotypic analysis across key stages of larval development through metamorphosis into reproductive adults (nonparasitic If) or parasitic juveniles (Iu); and 2) a population genomics study in which whole genomes will be sequenced for groups of each species from multiple streams where they are co-occur. These genomes will then be used to identify genomic regions that consistently differ between ecotypes. The selected student will lead all aspects of the project, including lamprey and sample collection, assessment of reproductive and trophic phenotypes, transcriptomic and genomic analyses, writing reports, and publishing results in scientific journals. Support and training will be provided by the PI (T Buchinger) and co-PIs at MSU (J Hume and Y-W Chung-Davidson), USGS (N Johnson), University of Manitoba (M Docker), and University of Glasgow (A Jacobs). The student may also have opportunities to develop other projects that leverage research facilities at HBBS and the Genomics and Chemical Ecology Laboratory at MSU (https://www.lilabmsu.com/). Qualifications: B.Sc. or M.Sc. (preferred) in biology, evolutionary biology, bioinformatics, genomics, or a related field. Experience or interest in fieldwork, morphology, and bioinformatics. Location: Primary location is MSU (East Lansing, MI) with regular sampling trips (~3 days each) to HBBS (Millersburg, MI), lamprey sampling throughout the Great Lakes region, and the opportunity for a visit (~3 months) to University of Glasgow (Glasgow, Scotland) to work with Co-PI Jacobs on analysis of genomics data. Start date: March-June 2026. Stipend: ~$35,000/ year; health insurance and tuition waiver. Funding is expected for 4.5 years. To Apply: Send a letter of interest including a brief summary of background and qualifications and a CV to Tyler Buchinger (buching6@msu.edu). Review of applications will begin 17 Nov 2025 and continue until the position is filled. Tyler Buchinger Assistant Professor Department of Fisheries and Wildlife Michigan State University Office: (517) 355-4106 Google Scholar profile "Buchinger, Tyler" (to subscribe/unsubscribe the EvolDir send mail to golding@mcmaster.ca)