Call for a PhD Position in Evolutionary Biology at Doñana Biological Station, CSIC , Seville, Spain Project: Timing of amphibian development: Processes, outcomes and links to ecology and evolution(TADPOLE) - 4-year fully funded contract for a doctoral candidate to obtain a PhD studying the evolution of developmental plasticity in spadefoot toads, combining fieldwork, experimental work with tadpoles, ecophysiology, and functional genomics. - Supervised by Dr. Ivan Gomez-Mestre, and joining the Ecology, Evolution and Development Group . - The contract has associated funding to cover short term stays at international R&D centers to train in further developing specific skills, taking advantage of a wide network of international collaborators. - Tuition fees for doctoral studies at a Spanish University will also be covered. Project description A major challenge in evolutionary biology is to understand how environmentally-induced changes in the regulation of development can evolve under selection into constitutive differences in adaptive traits. We will use the evolution of developmentally plasticity in spadefoot toads as a study system to understand the link between phenotypic plasticity and adaptive divergence across populations. This PhD project will focus specifically on the developmental plasticity of amphibian larvae in response to two main environmental factors: predation risk and risk of pond drying. It will be a highly integrative project, addressing fundamental aspects of phenotypic plasticity, such as adaptation to multiple environmental drivers, trade-offs underlying antagonistic responses, and plasticity costs. The project contemplates sampling spadefoot toad populations in different regions of Spain, including Doñana National Park, and experimentally modifying environmental conditions over the course of development. Using ecophysiological approaches and functional genomics, we will seek to understand the mechanisms allowing amphibian larvae to integrate environmental inputs and generating appropriate phenotypic responses, comparing populations that have diverged in their degree of responsiveness. Requisites - Master's degree in Biology (MSc) - High motivation to pursue a career in evolutionary biology - Advanced English speaking and writing skills - Experience in bioinformatic/genomic analyses will be valued When to apply Pre-selection of candidates is starting immediately, with position remaining open until filled. Official selection process likely to take place in October-November 2025, but candidates are strongly advised to submit expressions of interest by October 5th. Interested applicants should send the following documents to igmestre@ebd.csic.es: - CV, indicating two referral contacts (name and email address). - A motivation letter stating your interest in pursuing a scientific career and your interest and adequacy for this project in particular. Please refrain from using LLM-generated text, as we value genuine scientific motivation. Ivan (to subscribe/unsubscribe the EvolDir send mail to golding@mcmaster.ca)