The research group of Dr. Elena Jovanovska at the Department of Environmental Sciences, University of Basel is offering a fully funded, four-year (100%) PhD position in evolutionary biology of microalgae, as part of the SNSF Ambizione project: 'Key functional and morphological innovations in a rapid evolutionary radiation of diatoms in African Lake Tanganyika'. The project will investigate the evolutionary process of adaptive radiation and the associated key innovation of diatoms in African Lake Tanganyika. Project Description: Diatoms are among the most successful life forms on Earth, having taken over a substantial role in regulating the global oxygen balance and climate that other microalgae maintained for nearly a billion years. Their ecological and evolutionary success has made them dominant and diverse, yet the mechanisms behind this success remain poorly understood. This project seeks to address this question by examining the role of adaptive radiation and associated key innovations in the evolution of freshwater diatoms. By integrating morphology, physiology, genomics, transcriptomics, and computational modeling, we aim to (i) determine whether a newly discovered assemblage of diatoms of the genus Diploneis from Lake Tanganyika has diversified through adaptive radiation, and (ii) identify the morphological and physiological innovations underlying their rapid diversification. Your position: Your work will focus on physiological innovations related to sterols, with specific aims to (i) Identify sterol composition and distribution among species, (ii) detect sterol-related innovations and associated genes by examining genomes for signatures of positive selection, and (iii) explore sterol functions and their underlying regulatory genetic networks using comparative transcriptomics. Preferred Experience: - Diatom culturing and single-cell DNA isolation - Phylogenetics or genomics - Experimental and transcriptomic work - Sterol analysis - Fieldwork experience and readiness for multi-week sampling expeditions to remote sites in Tanzania, Zambia, and Malawi Qualifications: - Master's degree in biology, ecology, evolutionary biology, microbiology, physiology, genetics, or related field - Experience with diatom culturing, single-cell DNA isolation, phylogenetic, and sterol analyses is an advantage - Fluent written and spoken English - Willingness to conduct field research in remote areas of Africa Collaborations: The project involves national and international collaborations, primarily with Prof. Nemiah Ladd and Prof. Walter Salzburger (University of Basel), Prof. Andrew Alverson (University of Arkansas), and Prof. Cyprian Katongo (University of Zambia), as well as other partners who offer opportunities for field research, joint analysis, and networking. Application: Please submit a CV, a one-page motivation letter outlining your research interests, and two reference letters to elena.jovanovska@unibas.ch. Application Deadline: November 15, 2025 Starting Date: January 2026 More information and application details: https://jobs.unibas.ch/offene-stellen/phd-position-in-evolutionary-biology-of-microalgae/64cf91a6-b134-4bf9-a563-9efbf83ced7e For further inquiries, contact elena.jovanovska@unibas.ch Elena Salzburger (to subscribe/unsubscribe the EvolDir send mail to golding@mcmaster.ca)