We have a PhD position available in the University of Glasgow's College Futures Themes PhD Programme. This position is fully funded for stipend and tuition fees, and includes a research budget. The studentship is hosted at the University of Glasgow School of Biodiversity, One Health & Veterinary Medicine and supervised by Kathryn Elmer, Maureen Bain, and Neil Evans. Please find more information here https://www.gla.ac.uk/colleges/mvls/graduateschool/phd-research-opportunities/futures-programme/projects/fundamentals-of-life/fol22kathrynelmer/ The project is offered within the Future Themes programme, with final candidate selection handled by the programme committee. Project: FUNDAMENTAL MECHANISMS OF REPRODUCTION: VIVIPARITY IN AN EVOLUTIONARY CONTEXT Summary: Live-bearing reproduction (viviparity) is a key evolutionary innovation in amniotes, enabling extended maternal-foetal communication and influencing traits such as immune tolerance, nutrient transport, and birth timing. While viviparity is well-known in mammals, it has evolved independently in other lineages, making it difficult to reconstruct its origins. This project uses the European lizard Zootoca vivipara, a uniquely bimodal species with both egg-laying and live-bearing populations, to investigate the physiological and gene regulatory basis of viviparity. Its recent evolutionary transition and simple placental structure make it an ideal model for studying early pregnancy evolution. The research will compare reproductive tissues and gene expression between oviparous and viviparous individuals, quantify maternal-foetal communication using hormone assays, and explore foetal signalling through maternal tissue responses. These insights will advance understanding of reproductive trait evolution across amniotes. The student will gain training in molecular biology, next-generation sequencing, histology, microscopy, hormone analysis, and bioinformatics. They will also develop skills in animal welfare, data management, and scientific communication. Working within a collaborative and experienced research group, including international partners, the student will contribute to resolving fundamental questions in reproductive biology and evolutionary development. The student will join an active, collegial, and dynamic research group with on-going funded project in this topic. The successful candidate for this project is likely to be someone with a strong theoretical background in evolution, developmental biology and/or genetics, who can show evidence of practical laboratory and analytical experience in an appropriate field, and demonstrated aptitude for and interest in biology research. Experience with lizards and a drivers licence would be an asset. The University of Glasgow ranks in the world's top 100 universities. The School of Biodiversity, One Health & Veterinary Medicine is an outstanding research and teaching unit, with many opportunities for collaboration and discussion in a supportive and productive environment. Glasgow is a lively cultural city on the doorstep of the beautifully rugged Scottish Highlands. Start date: Oct 2026. Eligibility: This studentship is open to UK home students only. Deadline: Closing date for applications will be January 12th 2026 Informal inquiries to Kathryn Elmer in advance of the deadlines are welcome. Kathryn Elmer (to subscribe/unsubscribe the EvolDir send mail to golding@mcmaster.ca)